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Nardia & Solenostoma;

Jungermanniaceae                           PICTURE KEY TO NARDIA & JUNGERMANNIA

Scroll down for Nardia geoscyphus and Solenostoma gracillimum

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Nardia scalaris Gray                                                                                              Ladder Flapwort

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A small liverwort with shoots only c.1.5mm wide, with the two rows of incubous lateral leaves folded upwards like closed butterflies wings, with the front margins expanded so that they curve downwards beside the stem. They usually get larger towards the apex of the plant. The cells have 1-3 large smooth, glistening oil bodies that persist for several years on drying in a packet, in contract to the closely related N. geocyphyus that has dull granular oil bodies that disappear within 6 months of drying. It is very similar to Solenostoma gracillimum, but the genus Nardia differs in having underleaves. These are stiff, narrowly pointed, and usually stick out conspicuously, though they can sometimes be difficult to see against a mass of rhizoids. Most populations of Solenostoma gracillimum also have a conspicuous band of large pale cells bordering the underleaves, but some forms do not and can be confused with N. scalaris, necessitating a thorough search for the presence or absence of underleaves. Although dioecious, in contrast to the monoecious N. geosyphus, it is often fertile, and is further separated from N. geoscyphus by its rounded, unnotched leaves.

A plant of nutrient poor, mineral soils with a pH typically of less than 3. In Essex it is a rare plant of bare banks kept moist by overhanging vegetation, and is probably on the verge of extinction due to metalling of the  rides in Epping Forest, a decline in acid rain, and trampling. Despite the insertion of fibre optic cable involving cutting back the roadside bank of its most stable site on the south side of the Theydon Road just east of Jacks Hill Bog, it has recolonized the bank, sheltered by patches of Calluna. It is now only known from this one site. Geologically, the localities are largely confined to exposures of Bagshot Sands,. Woodford Gravels or Older Head.

All records:

TQ49

40,94

18

Buckhurst Hill, by Church Road Pond, 1967. Bernard Tavenor Ward. [on Woodford Gravel]. [possible a mistaken record for Solenostoma gracilimum or present as well?]

 

416,967

18

Epping Forest, dry patch by the Green Ride just east of Earl’s Path, (conspicuous underleaves) June1972. Kenneth J Adams. [Woodford Gravel]

 

43,99

18

Epping Forest, locally abundant by Theydon Road between Wake Arms and Theydon Bois, c.fr. 1962 and 1964. Eric Saunders.

 

435,997

18

Epping Forest, Jack's Hill,  along northern sector of Green Ride ditches, June 1968. Kenneth J Adams.

  4327,9942 18 Epping Forest, steep north facing roadside bank just east of Jacks Hill Bogs, with Lophozia ventricosa and Diplophyllum albicans. 15 December 1985. K.J.Adams.
  4327,9942 18 Epping Forest, steep north facing roadside bank just east of Jacks Hill Bogs, abundant, 1968; 29 July 1981. Kenneth J Adams. & 19 November 1994. Tim Pyner.

 

4327,9942

18

Epping Forest, steep north facing roadside bank just east of Jacks Hill Bogs, several patches among heather clumps with Diplophyllum albicans. KJA et Essex Bryology Group. 12 Feb 2011. [Bagshot Sands]

 

4309,9980

19

Epping Forest, St. Thomas’ Quarters, roadside opposite Wake Arms Bog North, with Gynocolea inflata. 28 March 1995. KJA. Still present 1996 & 1997 (now gone). [Older Head].

TL40

49,02

18

Colliers Hatch, track to Mount Wood, by the new plantation, September 1972. Stephen S Licence. [Older Head].

TQ69

61,91

18

Thorndon Park, 1958, Kenneth Alvin & Alan J Pettifer.

 

 

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Nardia geoscyphus (de Not.) Lindb.                                                           Earth-cup Flapwort

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TQ(51)49 41,98 18 Epping Forest, Little Monk Wood, pond-side by road (Little Wake Valley pond?). 1964. Eric Saunders.
  422,992 18 Epping Forest, Wake Road, roadside ditch by Sunshine Plain. March 1964. Eric Saunders.
  42-3,99 18 Epping Forest, nr. Wake Arms, damp sandy path to Long Running. c.fr. October 1962. Eric Saunders. Transactions British Bryological Society 4. p.484.
   c.430,992 18 Epping Forest, Jack's Hill, stream bank southTheydon Road, western tributary of Tippa Burn. 2 December 1967. Kenneth J Adams (herb.KJA).
TL(52)60 63-64,01 18 Mill Green, plentiful sides of gravel pit. September 1962. Eric Saunders.

 

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Solenostoma gracillimum (de Not.) Lindb.                                                                                Crenulated Flapwort

[Solenostoma crenulatum, Jungermannia gracillima, Plectocolea crenulata] __________________________________________________________________________________

 

All records:

 
TQ(51)49 481,941 18 Hainault Forest, on bare clay beside path, with Pellia epiphylla. 5 July 1967. Kenneth J Adams (herb.KJA)
  408,941 18 Buckhurst Hill, beside Church Road Pond, one patch c.30cm across growing vertically, 4 September 1967. Kenneth J Adams (herb.KJA).
  4187,9697 18 Epping Forest, the Green Ride, poached wet clay on north side, east of Earl's Path. 13 June 1971. KJA. (large marginal cells).  
  41,96-7 18 Epping Forest, The Green Ride, 1967. Kenneth J Adams.  
  424,979 18 Epping Forest, The Green Ride, in the bottom of Bellringers Hollow, south side of track. September 1967. Kenneth J Adams.
  4238,9793 18 Epping Forest, beside the Green Ride on west side as it rises west out of Bellringers Hollow, with Fossombronia wandraczekii. September 1969. and  9 October 1971. Kenneth J Adams.
  4244,9791 18 Epping Forest, beside the Green Ride on west side as it rises west out of Bellringers Hollow, 15 October 1980; KJA; with Fossombronia wandraczekii. 29 July 1981. Kenneth J Adams &              Monica Milne-Smith.
  426,993 18 Epping Forest, bank of gravel working just south of Wake Arms roundabout. 6 March 1993. Kenneth J Adams
  43,99 18 Epping Forest, Oak Hill, The Ditches Ride. 1967. K.J.Adams.
  431,997 18 Epping Forest, poached horse ride by Epping Road opposite Lodge Road Junction, plentiful in wet mud.15 September 1978. Kenneth J.Adams.
  429,996 &

431997

18 Epping Forest, large patch midway between Wake Arms and Lodge Road turn off, and two smaller patches opposite Lodge Road turn off, poached horse ride parallel to east side of Epping Road. 20 August 1986. Kenneth J.Adams.
TL(52)40 49,02 18 Stanford Rivers, High Wood. date? Eric Saunders
  470,036 19 Epping Lower Forest, stream bank edge of ride with Calypogeia fissa. 12 August 1978. Kenneth J Adams
  476,035 19 Epping Lower Forest, east side of N-S track, 24 March 1978.  
  49,02/50,02 18 Ongar Park Wood. 1964. Eric Saunders
TL(52)52 515,296 19 Quendon Wood, plentiful with Riccardia chamedryfolia and Scapania irrigua, wet clay woodland ride. 18 July 1967. Still there in main E-W ride 10 April 1983. Kenneth J. Adams et EFC. (herb.KJA.)
      Quendon Wood, still there in main E-W ride 10 April 1983. Kenneth J. Adams et EFC
TQ(51)69 61,91 18 Thorndon Park North ???  
  686,951 18 Billericay, Norsey Wood, on Clay. var. gracillima, in base rich hollow in angle between the first fork in the stream, with Plagiochila aspenlioides. 10 September 1972. Kenneth J Adams.(herb.KJA).
TL(52)81 88,14 19 Tiptree Heath, 1862 Ezekiel Varenne
TL(52)82 87,28 19 Chalkney Wood, 1956. Francis Rose & Edward C Wallace.
TL(52)92 96,28 19 West Bergholt Heath, var. gracillima. damp peaty ground. 1957. Francis Rose

Trans. Brit.Bryol.Soc. 4. p.152.1961

TM(62)12 1134,2458 to 1140,2460 19 Little Bentley Hall Wood, 1969 and 10 August 1973. Kenneth J. Adams & A. J Harrington.
  158,209 19 Weeley Hall Wood, marshy cart track, east west central ride, c.half way along. 26 May 1986. Kenneth J Adams.
TM(62)13 18,31 19 Stour Wood, 1951. Frances Rose