Dryopteris carthusiana

ESSEX BOTANY AND MYCOLOGY GROUPS

Dryopteris carthusiana:

Scroll down for records.

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Dryopteris carthusiana (Vill.) H.P.Fuchs Narrow Buckler-Fern

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Light 6. Wetness 8. pH 4 to 6. Fertility 4. Height <1m

Essex Status: Native.

Native. This fern is frequently confused with the much commoner D. dilatata. It grows in a wide range of damp lowland habitats where the atmosphere is permanently humid and the soil moderately acidic in reaction (pH4-6); wet heath, wet woodland, stream sides, fens, bogs and lake margins, preferring deep rich alluvial soils rather than acid boggy but infertile sites. It frequently grows in water-saturated Sphagnum and Juncus dominated communities, under relatively open-canopy, Alnus, Salix or Betula carr. Its distribution is thinning out nationally through loss of habitat. The fronds are generally narrower, paler-green and less leathery than those of D. dilatata, and are generally held erect, only flopping over at the tip; and emerge from a creeping rhizome, so that bunches of fronds appear to arise in lines from their typically buried crowns, rather than flopping over in an arc, and arising from an obvious single shuttlecock as in D. dilatata. The basal stipes are sparcely clothed with delicate pale-brown, soft, concolorous scales, in contrast to the usually centrally dark-brown-striped rather stiff scales of D. dilatata. As in D. dilatata the fronds are divided 3-times (unlike the D. filix-mass-affinis ferns) into pinnae, then into pinnules and finally into serrated-margined pinnulets, the saw-like teeth incurving and ending in needle-like spines visible with the naked eye. D. dilatata frequently grows amongst it, though elevated on less water-soaked rotting tree trunks, tree bases or hummocks, and they hybridise, the offspring [D. x deweveri] resembling a narrow-fronded D. dilatata with dark scales and largely abortive spores. The latter is under-recorded in Essex. D. carthusiana is thinly scattered across the county, largely absent in the north west sector, but quite common in damp marshy areas and along rides in woods. Very much under-recorded.

Post 1930 records:

TL(52)30	39,03	19	Galleyhill Wood. Flora of Essex 1974.
TQ(51)49	408,981	18	Epping Forest, Rat's Lane stream, boggy patch, 2-3 small plants. 8 September. 2010. Kennth J Adams.
	409,981		Epping Forest, small woodland pool south east of Rat's Lane, 2 small plants. 8 September. 2010. Kenneth J Adams.
	41,99	18	Epping Forest, by Claypit Hill, single small plant in deep ditch. 21 May 2009. Kenneth J Adams.
	41,96	18	Epping Forest, Loughton Brook just south of Baldwins pond. 1973. Kenneth J Adams.
	422,970	18	Epping Forest, Woodberry Hollow. Flora of Essex 1974.
	434,994	18	Epping Forest, Jack's Hill Bog South. 28 July 1993. Amanda Samuels.
	402,954	18	Epping Forest, [area to NW of Connaught Waters] 1993-1994. Amanda Samuels.
	402,947	18	Epping Forest, damp scrub area to west of Ching Brook, just south of Rangers Road. 2 May 2009. Kenneth J Adams et Essex Bryology Group.
	417,970	18	Epping Forest, Honey Lane. 15 September 1996. Tim Pyner.
	418,994	18	Epping Forest, Woodredon Hill, single plant, stream bank, just south of the bend in the road. 17 October 2009. Tim Pyner et Essex Bryology Group.
	418,974	18	Epping Forest, 1-2 clumps, stream draining Loughton Camp. 1996. Amanda Samuels.
	4361,9952	18	Epping Forest, several small clumps in Oak Hill Main Bog. 8 September 2010. Kenneth J Adams.
	4363,9938	18	Epping Forest, 1-2 clumps in boggy section of Oak Hill stream. 8 September 2010. Kenneth J Adams.
	4359,9919	18	Epping Forest, 3+ clumps in boggy flush, second main stream south from Oak Hill car park to east of Green Ride. 8 September 2010. Kenneth J Adams.
	43307,99332	18	Epping Forest, Tipper Burn Bog, single medium sized plant. September 2010. Kenneth J. Adams.
	4308,9993	19	Epping Forest, c.4-5 small plants in boggy patch half way down stream draining Lodge Rod Bog South. 22 October 2010. Kenneth J.Adams.
	4308,9985	19	Epping Forest, several plants under bracken in spring head bog just below Lodge Road car park. 22 October 2010. Kenneth J.Adams.
TL(52)40	431,000	19	Epping Forest, Lodge Road Bog North, in the Sphagnum mire at the inlet end. 1953. K J Adams.
	431,000	19	Epping Forest, Lodge Road Bog North, 1995. Tim Pyner.
	431,000	19	Epping Forest, Lodge Road Bog North, c, 50 clumps, in the Sphagnum mire at the inlet end. May 2006 and October 2010. K.J.Adams.
	4304,000	19	Epping Forest, c.10 plants in boggy patch half way down stream draining Lodge Road Bog North. 22 October 2010. Kenneth J. Adams.
	4374,0023	18	Epping Forest, 5 small plants in Ambersbury Banks bog, north of Green Ride. September 2010. Kenneth J. Adams.
	4378,0016	18	Epping Forest, c.6 medium plants in perched bog just south of the Green Ride. September 2010. Kenneth J. Adams.
	44,07	19	Harlow, Parndon Wood north, with Athyrium opposite the centre building. 10 June 2009. Kenneth J Adams et EFC.
	44,06	19	Harlow, Parndon Wood south, abundant along upper reaches of stream above bridge. 10 June 2009. Kenneth J Adams et EFC.
	47,03	19	Epping Lower Forest. Flora of Essex 1974.
	472,034	19	Epping Lower Forest, [east of the lake] July 2000. Amanda Samuels.
	48,02	18	Gernon Bushes, N.R. small quantity. 16 June 2007. Tim Pyner.
TL(52)41	49,17	19	Wallbury Dells. Flora of Essex. 1974.
TL(52)43	43,36 or 43,35	19	Roughway Wood. Flora of Essex 1974.
TQ(51)58	57,85	18	Cranham Marsh. Spring Wood. 16 June 1965. Arthur Adams.
TQ(51)59	50,98	18	Stapleford Tawney, Shales More, marshy flushes, scattered. 8 April 1995,. Tim Pyner.
	591,911	18	Warley, Holdens Wood. 1981. Kenneth J Adams.
	561,965	18	Coxtie Green, The Mores, plentiful under Sambucus. 14 July 1960. Kenneth J Adams.
	51,92	18	Bedford's Park. Flora of Essex 1974.
	5129,9707 & 5134,9705	18	Curtismill Green, old gravel workings, 2 clumps in north west working,c.10 clumps large southern working, 29 August 2010. K.J.Adams.
	569,952	18	South Weald Park. 12 September 1971. Martin Rickard; & Flora of Essex 1974.
	573,991	18	Kelvedon Hatch, The Coppice, large clump with D. x deweveri, under Alder. 25 May 2003.Tim Pyner et EFC. Still there 19 September 2015. K.J.Adams et EFC.
	5829,9107	18	Warley Place, several clumps by NE pond, 13 July 2016. Ken Adams & Tim Pyner.
TL(52)50	57,04+	19	Norton Mandeville. Flora of Essex 1974.
TL(52)51	53,11	19	Matching Green, 29 September 2003. Shirley Watson.
	5416,1996	19	Hatfield Forest, Gravel pit Coppice, one small plant near edge of coppice with one P. setiferum and one Athyrium under old Sambucus. 21st September 2013. Pteridological Society.
	5433,1998	19	Hatfield Forest, Gravel Pit Coppice, in damp hollow one large convincing plant with c.10 Dryopteris affinis. 21st September. 2013. Pteridological Society.
TL(52)52	5428,2009	19	Hatfield Forest, Gravel Pit Coppice, one large plant nr NE edge. 21st September. 2013. Pteridological Society.
	5427.2010	19	Hatfield Forest, Gravel Pit Coppice, second large plant nr NE edge. 21st September. 2013. Pteridological Society.
TL(52)54	57,41	19	Shadwell Wood. Flora of Essex 1974.
TQ(51)69	60,91	18	Brentwood, Kent's Wood, marshy areas, scattered. 13 January 1995. Tim Pyner.
	60,92	18	Brentwood, Harts Wood, frequent in places. 28August 1984. Tim Pyner.
	61,90-61,91	18	Brentwood, Thorndon Park North, scattered in marshy area. 6 November. 2004. Tim Pyner.
	61,93	18	Brentwood, Thrift Wood, in ponds and boggy areas. 27 December 1986. Tim Pyner.
	655,985	18	Ingatestone Hall, Wood, single plant. 19 July 1998. Graham Smith.
	68,95-69,95	18	Billericay, Norsey Wood, frequent. 7 September 1997. Tim Pyner et EFC.
	69,95	18	Billericay, Norsey Wood east, patch by middle pond. 15 July 2009. Kenneth J Adams.
	68,95	18	Billericay, Norsey Wood, abundant all over the wood in damp and apparently dry areas. 15 July 2009. Kenneth J Adams.
TL(52)60	60,04	19	Norton Heath, banks of old gravel workings. Flora of Essex 1974; Still there 7 October 1991. Tim Pyner.
	630,026	18	Birch Spring Wood. c.1970. Martin Rickard. & Flora of Essex 1974.
	629,025	18	Highwood, Barrow Wood, a few along stream. 2002-2006. Graham Smith.
	636,019	18	Ingatestone, The Mores, Stonymore Wood. 30+ in coppiced area.
	63,02	18	Highwood, Deerslade Wood. damp areas 15-20 plants. 7 July 2006. Graham Smith.
	64,02	18	Highwood, Ellis Wood, several along stream.29 May 2002. Graham Smith.
	69,02	18	Galleywood Common, scattered in boggy areas. 1988-2004. Tim Pyner.
	654,042	18	Highwood, Edney Common, 2 plants by stream 2002. Graham Smith.
	663,041	18	Highwood, Edney Common, single plant. 1999. 2 plants by stream 2002. Graham Smith.
	666,033	18	Writtle Park Wood, a few by stream. 9 June 2000. Graham Smith.
	675,043	18	Writtle, South Wood. 2 plants. 2000. Graham Smith.
TL(52)61	63,18	19	Great Dunmow, Garnets Wood, scarce marshy areas. 29 April 1995. Tim Pyner.
	63,18	19	High Easter, Barnston Lays. Flora of Essex 1974.
TQ(51)78	?????	18	Thundersley Glen. Flora of Essex 1974.
TQ(51)79	704,978	18	Stock, by Kiln Common, frequent around spring, scattered elsewhere. 30 November 1997. Tim Pyner.
TL(52)70	772,068	18	Little Baddow, Blakes Wood. 20 June 1969. Martin Rickard. Flora of Essex 1974.
	77,06	18	Little Baddow, Blakes Wood, frequent in marshy areas. 24 May 2008. Tim Pyner.
	772,068	18	Little Baddow, Blakes Wood, 20 May 1969. Martin Rickard.
	770,068	18	Little Baddow, Blakes Wood, 2 plants on shelf north of stream. Tim Pyner et Wildflower Group 10 April 2014.
	772,062	18	Little Baddow, Blakes Wood, 1 plant marshy area at stream junction. Tim Pyner et Wildflower Group 10 April 2014.
	78,06	18	Little Baddow, Poors Piece, 16 July 2005. Kenneth J Adams et EFC.
	78,06	18	Little Baddow, Pheasanthouse Wood Bog, Kenneth J Adams et EFC.
	78,06	18	Little Baddow, Woodham Walter Common Bog. Kenneth J Adams et EFC.
TL(52)71	74,16-74,17	19	Great Leighs. Mann Wood, scattered, marshy slopes southern end of wood. 1 May 1992. Tim Pyner.
TL(52)73	78,31-79,31	19	Sible Hedingham, Broaks Wood, scattered locally frequent. 20 May 1995 & 20 June 2009. Tim Pyner.
TQ(51)88	81,87	19	Leigh on Sea, Belfairs N.R. marshy areas by stream. 21 February 2002. Tim Pyner.
	81,88	18	Daws Heath. Flora of Essex 1974.
TQ(51)89	847,908	18	Hawkwell, Potash Wood, in boggy pool. Tim Pyner.
TL(52)80	804,052	18	Woodham Mortimer, Thrift Wood, marshy areas. 8 October 1989. Tim Pyner.
	80,05	18	Woodham Mortimer, Thrift Wood, margin of pond. 25 November 1990. Tim Pyner.
	80,05	18	Woodham Mortimer, Thrift Wood, 3-4 patches in recently coppiced area. 31 May 1997. Richard Carter.
	81,07	18	Woodham Walter, marshy areas south of Whitehouse Pit, scattered plants. 27 December 1997. Tim Pyner.
TL(52)81	87,16	19	Tiptree. 1986. Peter Wilson.
	88,14	19	Tiptree Heath, 1977. Colchester N.H.Museum. & 1987. Tiptree Management Committee.
	89,17	19	Tiptree. 1988. Peter Wilson.
	89,18	19	Tiptree/Messing cum Inworth. 1988. Peter Wilson.
	84,13+	19	Wickham Bishops, Chantrey Wood. Flora of Essex 1974.
TL(52)82	80,26	19	Stisted, Belchers Wood, a few plants, damp woodland. 26 June 1993. Tim Pyner.
	809,267	19	Stisted, Belchers Wood, patch with c.6 fronds. 26 June 1993. Jeremy Ison.
	87,27- 87,28	19	Earl's Colne, Chalkney Wood, frequent 1086-2005. Tim Pyner.
	871,274	19	Earl's Colne, Chalkney Wood, 2 clumps 21 August 1991. Jeremy Heath and Terri Tarpey.
	874,280	19	Earl's Colne, Chalkney Wood, 2 plants in wet flush. 14 May 1994. Jeremy Ison.
	875,281	19	Earl's Colne, Chalkney Wood, scattered plants. 14 May 1994. Jeremy Ison.
TL(52)83	88,35	19	Essex Wildlife Trust Records. May 1984.
	873,355	19	Alphamstone Gravel pits, a few plants, marshy area by stream. 3 July 1996. Tim Pyner.
	854,336	19	Pebmarsh, [marshy woodland west of Cross End]25 September 191997. Nigel Cooper.
	869,361	19	Alphamstone, The Ashground. 1968. Kenneth J Adams & Stanley T Jermyn; and 4 January 1970. Martin Rickard.
	87,36	19	Great Henny, Loshes Brook. 1968. Kenneth J Adams & Stanley T Jermyn.
TQ(51)99	93,96	18	Creeksea Place. Flora of Essex 1974.
TL(52)91	90,17	19	Tiptree, Pods Wood. 1989. Peter Wilson
TL(52)92	904,264	19	Hoe Wood. 1 plant. May 1995. David Baker.
	92,26	19	[Colne Valley] 5 June 1983. Jeremy Heath, Terri Tarpey and Alan Wake.
	930,268	19	Aldham C,P., wood near Fidler's Farm. 20 June 1971. Martin Rickard,
	944,265	19	Fordham Heath, 29 July 1997. CNHS;1998. CNHS.
	944,264	19	Fordham Heath, a few plants by side of ditch downstream from well. 30 July 1996. CNHS.
	94,28	19	West Bergholt, Hill House Wood, frequent in marshy areas. 31 October 1994. Tim Pyner.
	947,281	19	West Bergholt, Hill House Wood, by stream 1994. CNHS.
	964,290	19	Great Horkesley, Pitchbury Wood, 1 plant in damp area. 26 May 1997. Anne Guiver & Terri Tarpey.
	963,294	19	Great Horkesley, Pitchbury Wood, 1 plant edge of large pond. 8 June 1997. Anne Guiver, Terri Tarpey & Tim Pyner.
	967,204	19	Layer de la Haye, Heather Fields, 30 June 1992. CNHS.
	968,208	19	Layer de la Haye, Heather Fields, straggly patch in wide damp ride, October 2010.Tim Pyner et Essex Bryology Group.
	98,20	19	MOD Berechurch, Friday Wood, scattered in marshy areas. 6 November 2004. Tim Pyner.
	998,214	19	MOD Berechurch. 10 July 2007. Jeremy Heath & Terri Tarpey.
	999,212	19	MOD Berechurch. 10 July 2007. Jeremy Heath & Terri Tarpey.
TM(62)01	04,19	19	Fingringhoe Wick N.R. 1989. Laurie Forsyth; scarce, marshy areas. 20 July 1991. Tim Pyner.
TM(62)02	009,222	19	MOD Middlewick & Doneyland, 19 February 2007.Terri Tarpey & Jeremy Heath.
	017,219	19	MOD Middlewick & Doneyland, 28 July 2003. Terri Tarpey & Jeremy Heath.
	019,211	19	East Donyland Heath, a few plants in ponds. 14 November 1992. Tim Pyner; Birch Brook, 2003. Jeremy Heath & Terri Tarpey.
	019,219	19	East Donyland Heath, c. 10 plants. 8 August 1995. Terri Tarpey & Jeremy Heath.
	025,264	19	Welsh Wood. 11 June 1996. CNHS.
	03,21	19	1987. Chris Gibson.
TM(62)03	01,32	19	Langham, by Black Brook, 1 plant seen in marshy area. 26 October 1998. Tim Pyner.
	07,31	19	Boxted-Langham, by stream. Stanley Jermyn; Humberlands,
	07,31	19	Lawford, Shir Burn, Alder carr, frequent. 15 November 1995. Tim Pyner.
TM(62)11	127,180	19	St. Osyth, Riddles Wood. 1 plant by ride. 30 May 1993.CNHS field trip.
	14,19	19	Malden Wood. Flora of Essex 1974.
TM(62)12	11,24	19	1961. CNHMus.
	10,24	19	Ravens Green. Flora of Essex 1974.
	16,21	19	Weeleyhall Wood. 11 May 1999. CNHS.
	157,210	19	Weeleyhall Wood. 17 November 1995. Terri Tarpey.
TM(62)13	18,30	19	Stour Estuary. 1985. Russell Leavett.
	189,310	19	Ramsey, Stour Wood, 1 plant. 15 October 1995. Terri Tarpey; 1997. Russell Leavett.
	19,30	19	Stour Estuary. 1985. Russell Leavett.
	18,31-19,31	19	Ramsey, Stour Wood, scattered along damp rides. 1991-1999. Tim Pyner.
	1932,3129 & 1934,3130	19	Ramsey, Stour Wood, two definite linear clumps either side of open ride. 9 October 2010. K.J.Adams.
TM(62)22	214,273	19	Bramble Island, edge of borrow dyke. 1990. National Rivers Authority survey,
TM(62)23	20,31	19	Copperas Wood, small colony just south of railway line. 1983. Tim Pyner; 1999. Russell Leavett.
	21,31	19	Stour Estuary. 1991. Russell Leavett.

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Dryopteris x deweveri (J.T.Jansen) Jansen & Wacht. (D. carthusiana x dilatata )

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This hybrid is overlooked nationally as well as in Essex, but is believed to be common where the two parents grow together.

It has been noted at Kelvedon Hatch by Tim Pyner (specimen deposited in Southend Central Museum Herbarium), and as a record to be confirmed: by the Ching Brook, just South of Ranger's Road in Epping Forest.

Danbury, Scrub's Wood, in axil of stream junction, TL79002,06154. 25 July 2015. Spores aborted. Tim Pyner et British Pteridological Soc.